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been syndactylous, and the authors of the accountl of these bones
think that the fourth toe may have shared in this syndactyly.
The metatarsal of the fifth digit is enormously expanded at its

FIG. 73.óDi-protodon australis*     (After Owen.)

and seems to have furnished a strong support to the
creature ; this is also seen in the metaearpal of the fore-limb.
Probably, therefore, Diprotodon was quadrupedal in its mode of
progression, with the em-
phasis laid upon the
little finger and the
little toe instead of, as
in ourselves, the first
toe The hind-foot of
the Diprotodon could
not be more unlike that
of a Kangaroo than it
actually is.

Another giant among
these Marsupials was the
genus Thylaeoleo, whose name was given to it by Sir Bichard
Owen on the view that it was a Marsupial Tiger, Sir W. Flower
has, however, controverted this opinion, and the genus is in fact,
in spite of its large size, closely allied to the Phalangers and

1 Stirling and Zietz, Mem. Roy. Soc. goudh Australia, i,;  see also a notice in
Nature, January 18, 1900.

FIG. 74.-

-Thylacoleo carnifesc.    Side view of skull,
(After Flower.)